Member Of Technical Staff (AI Security Engineer)
Perplexity is seeking a highly skilled, experienced, and hands-on AI Security Engineer to join our security team, driving the protection of next-generation AI systems against adversarial threats. In this role, you’ll design and implement robust mechanisms to secure self-hosted models, LLM APIs, agents, MCPs, and the co
What this role actually needs.
Perplexity is seeking a highly skilled, experienced, and hands-on AI Security Engineer to join our security team, driving the protection of next-generation AI systems against adversarial threats. In this role, you’ll design and implement robust mechanisms to secure self-hosted models, LLM APIs, agents, MCPs, and the co Responsibilities: - Define, build, and refine mechanisms to secure AI systems (including self-hosted models, LLM APIs, agents, MCPs, and other core components of the AI stack) against adversarial behavior of all kinds - Understand technically complex AI systems, identify potential weaknesses in their architecture, and implement improvements - At least 50% of time performing hands-on remediation. Also working closely with peer engineers to drive remediations - Plan and carry out threat modeling activities and realistic threat simulations across our offerings - Conduct cybersecurity evaluations and lead AI security assessments in a cross-functional environment - Develop initiatives that improve our capabilities to effectively evaluate AI systems and enhance the organization's prevention, detection, response, and threat hunting capabilities Requirements: - Hands-on coding and prompting experience. - Bachelor of Science or Master of Science in Computer Science or a related field, or equivalent experience - Be a technical and process subject matter expert regarding AI security services and attacker tactics, techniques, and procedures - Good understanding of LLMs, AI architecture patterns, machine learning models, and related technologies such as MCP - Understanding of application security principles and secure coding practices - Experience developing and implementing security procedures and policies Company context: Perplexity is an answer engine — an AI-native search product used by millions of professionals daily.
Day-to-day expectations
Perplexity lists these responsibilities for the Member Of Technical Staff (AI Security Engineer) role.
- Define, build, and refine mechanisms to secure AI systems (including self-hosted models, LLM APIs, agents, MCPs, and other core components of the AI stack) against adversarial behavior of all kinds
- Understand technically complex AI systems, identify potential weaknesses in their architecture, and implement improvements
- At least 50% of time performing hands-on remediation. Also working closely with peer engineers to drive remediations
- Plan and carry out threat modeling activities and realistic threat simulations across our offerings
- Conduct cybersecurity evaluations and lead AI security assessments in a cross-functional environment
- Develop initiatives that improve our capabilities to effectively evaluate AI systems and enhance the organization's prevention, detection, response, and threat hunting capabilities
What a strong candidate brings
These requirements are extracted from the source listing and normalized for UpJobz readers.
- Hands-on coding and prompting experience.
- Bachelor of Science or Master of Science in Computer Science or a related field, or equivalent experience
- Be a technical and process subject matter expert regarding AI security services and attacker tactics, techniques, and procedures
- Good understanding of LLMs, AI architecture patterns, machine learning models, and related technologies such as MCP
- Understanding of application security principles and secure coding practices
- Experience developing and implementing security procedures and policies
Why this listing is more than a copied job post.
Member Of Technical Staff (AI Security Engineer) is framed against UpJobz source checks, country scope, compensation visibility, and work-authorization signals so candidates can make a faster go/no-go decision.
United States tech market
United States roles on UpJobz are filtered for high-tech relevance, source freshness, and actionable employer detail before they are allowed into SEO surfaces.
Compensation read
$220K - $405K is visible before the click, so candidates can compare the role against local market expectations before applying.
Work authorization read
Current extracted signal: Open to TN, H-1B, and OPT candidates already in the United States. UpJobz treats this as a search signal, not legal advice, and links visa-sensitive roles back to the relevant visa hub where possible.
Location read
On-site roles in San Francisco should be compared against commute, local salary bands, and nearby employer demand.
Browse similar jobs
Turn this listing into an application plan.
This is the first pass at the premium UpJobz layer: a fast brief that helps serious applicants move with more clarity.
Next moves
- Tailor your resume around ai and llm instead of sending a generic application.
- Use the first two bullets of your application to connect your background directly to member of technical staff (ai security engineer) is a high-signal on-site role in san francisco, and it is most realistic for open to tn, h-1b, and opt candidates already in the united states.
- Open the role quickly if it fits and bookmark three similar jobs before you leave the page.
Interview themes
Watchouts
- $220K - $405K is visible, so calibrate your application around the posted range.
- Use open to tn, h-1b, and opt candidates already in the united states as part of your positioning so the recruiter does not have to infer it.
- Show concrete examples of succeeding in on-site environments.
Keywords to match against your background
Use these terms to decide whether your resume, portfolio, and recent projects line up with the role.
Apply through the employer source
Open the source listing from jobs.ashbyhq.com, confirm the role is still active, then apply on the employer or ATS page.
Source: jobs.ashbyhq.com · Source ID: cdbf6ccb-2078-4499-b0a6-af8a04754eee · Confidence: 94/100 · Last checked: May 7, 2026
How UpJobz verifies job sourcesContinue browsing tech jobs